I don't know what specifically will trigger this alert. I'm afraid it can be a lot of different things. I get this alert regularly, even with 128GB RAM and a 2TB ssd disk cache. There can be lots of causes, certain (combination of) effects, plug-ins, expressions, maybe a certain font used, other (adobe) apps running at the same time, complexity of the comp, using MFR, duration of uptime After Effects and/or computer, etc. Restarting AE will usually suppress the warning for a while. I would guess if you get this warning on a system with enough RAM available (32GB or more) there is some sort of memory leak going on in some process, filling up RAM, choking AE and it's memory management.
No idea what changed between 23 and 24 that would change it for your projects, I had this warning also regularly using 23... My experience with this warning is also not consistent enough to pinpoint it to certain effects/plug-ins/etc. One day I'll have this warning, then I don't see it again for the same project while it gets more complex.
It's like AE is walking a tight RAM rope, sometimes getting severely out of balance, slowing down or even stalling all progress, and sometime just tripping up completely, leaving it hanging on it's crotch...
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