This is caused by inconsistent color management between machines.
Hi Rick! Thanks for responding!
The thing is that I'm doing this test on a single machine that has both versions of After Effects installed on it....
Also is there a reason that one version would say "Millions" while the other says "Trillions"?
You can set the options for the color picker display in the Info Panel.
Any setting other than Auto Color Display will override how the values displayed in any color picker popup panel When the Info Panel is set to Auto the values in the color picker are driven by the Color settings in Project Preferences (Shift + Alt/Option + Ctrl/Cmnd + k).
Any expressions you use for color require a 4 digit RGBA array and the values are always decimal (float/32 bit). White would be [1, 1, 1, 1]. Red with a 50% transparency setting would be [1, 0, 0, .5]. Not all editable color properties in After Effects support alpha channels but a four-digit array is still the way to go unless it throws an error.
I hope this clears things up.