Google Play - Failed to verify JAR signature META-INF/CERT.RSA

I'm trying to update an Android app using Animate and 33.0.2.246 or 33.0.2.330.

The APK publishes fine, but when I upload I get this:

You uploaded an APK with an invalid signature (learn more about signing). Error from apksigner: ERROR (Jar signer CERT.RSA): Failed to verify JAR signature META-INF/CERT.RSA against META-INF/CERT.SF: java.security.SignatureException: Signing certificate has unsupported critical extensions

Any idea what the issue might be?