No, the changes to your path aren't due to the non-zero winding filling rule. That applies to how a fill (as opposed to a stroke) is determined based on how a path overlaps itself. What's happening with a paint brush stroke is that the intersection is telling Animate that despite how you've drawn the line, the there's a start and end to your path at that intersection. It's especially noticeable with the variable width profile. If you want to use a vector art brush and get an effect more like the default brush, look for a brush that doesn't have as noticeable of a tapered end like Chalk - Blunt. If you want tapered ends on the points where you do start and stop your path, draw the path with the uniform width profile and adjust the ends afterwards with the Width tool ( U).
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