PDF will export from Framemaker, but bookmarks and cross-references don't work

Hello - I’m having trouble with the bookmarks and cross-references in a PDF export from Framemaker. Once I create the PDF, none of the bookmarks and cross-references work. I’ve tried Publish to PDF and Save to PDF with no luck either way. I’ve also exported just one file to see if the links would work if I created one PDF at a time but that doesn’t work either. 

My process: Update book, Save all files in book, save book, Publish to PDF or Save to PDF.

I’ve included images of the following: PDF output settings, Settings.sts, list of book files.

Any help you can offer would be greatly appreciated. Thank you!