Right leg and both arms will not weld to torso (left leg works) — ACA 24.5 macOS

I am rigging a custom puppet in Adobe Character Animator.

My setup:
The Body group contains Torso, Left Arm, Right Arm, Head, Left Legs, and Right Legs.
The limb groups (Left Arm, Right Arm, Left Legs, Right Legs) are set as Independent and set to Attach To Body with Attach Style Weld.
Hip and Shoulder handles are placed at the seams.
Thigh layers attach to the Hip handles. Upper Arm layers attach to the Shoulder handles.

The left leg attaches correctly to the torso.
The right leg and both arms stay detached with a visible gap, even though the settings and handle placement match the left leg.

Questions:

1. What could cause one side to weld correctly while the other side does not?

2. Could mesh islands, origin placement, or the “Connect Each Island To” setting cause this problem?

3. Should Shoulder and Hip handles be placed on the child layers (Upper Arm or Thigh) or on the group layers?

Environment:
Adobe Character Animator 2025 (24.5)
macOS Sonoma
Puppet built from PSD

Steps to reproduce:

1. Import PSD with the structure above.

2. Mark limb groups Independent and set Attach To Body with Weld.

3. Add Left and Right Hip handles at the thigh seams and Left and Right Shoulder handles at the armpit seams.

4. Add Knee, Ankle, Wrist, and Elbow handles.

5. In Record mode, drag limbs or apply Physics. The left leg behaves correctly, but the right leg and both arms separate from the torso.

Expected: All limbs should weld to the torso at their hip or shoulder seams.
Actual: Right leg and both arms leave a gap from the torso.

I can provide screenshots of layer hierarchy and properties for the Left and Right limbs, as well as the PSD or exported puppet if needed.

Thanks for any guidance.