Following update to 25.1, AE asks me to save my current work-in-progess project every time I open it

I'm working on a Premiere project that I started in 24.x and because Media Encoder was updated to 25.1, the dynamic linking broke and I had to update Premiere as well, and then AE.

So now everything's on 25.1, and so every time I go back and use Edit Original in Premiere to open the AE composition, once AE has opened it asks me to save the AE project. It then forgets that it's saved that project, so it keeps opening the original converted one.

As an example, I've just opened the project "Project (converted).aep" using Edit Original and because it demanded I save it, I saved it to "Project (converted) v3.aep".

If I now close AE, when I reopen the project it will reopen "Project (converted).aep" and not "Project (converted) v3.aep" - and ask me to save it.

I've tried resetting AE's preferences, and it hasn't fixed it. Any ideas?