Frustrating issue with trim paths

I looked at your original file. As I expected, there were Groups on every layer I checked. I suspect that there were Groups on every layer. I also checked the graphics using the View/Outlines menu and saw that every shape was a closed and filled path. All of the Groups fouled up the XD - After Effects workflow, and converting the vector paths to shape layers will not fix the problem either.

The simplest workflow is to select all layers, lock them all, turn them all off, and then turn on layer 1. Add a new Track Matte layer above Layer 1 and give it a name. Then, decide how you want to reveal that layer and use the pen tool to draw a path with a wide enough stroke to hide the original path. When I started, I was ungrouping the layers, but then I realized that nothing was being gained. 

Here's an animated gif of the workflow required to create track mattes for the first dozen layers. Unless you want to group several layers and reveal them all with one track matte, you only have about a hundred to go. I compressed about 30 or 40 minutes of work into about 40 seconds.

When you have made all of the track mattes, open the AI file as a composition, retaining layer sizes, and then convert only the Track Matte layers to shape layers. 

It would probably take about the same time to create the shape layer track mattes in After Effects, but I think I'm a little faster at creating complex paths in Illustrator than in After Effects. Unfortunately, unless the artwork is remade with animating using After Effects in mind, the project will take a lot of time to set up.

First, Trim Paths only works with Shape Layers. Second, anything but a simple stroke in Illustrator will convert to a filled closed path, so you cannot use brushes or taper the paths in Illustrator.

It looks like you have converted all the vector layers to shape layers, but I cannot tell how cleanly the conversion worked. Select the layers, press "uu" to reveal all modified properties, and start digging into the shape of the layer structure.

Another option for working with complex Illustrator files would be to install Adobe XD. You can use XD to create a new file, then go to the File/Import menu and import your Illustrator file. Then, without doing anything else, create an After Effects project file using the File/Export/After Effects menu. Open the file, and the comp will be filled with much simpler and cleaner shape layers instead of embedded Illustrator layers. I use this workflow all the time.