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Hi Community!
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The most likely cause is your inconsistent use of the collapse transformations options that may affect render order. To debug the situation you would need to turn it off in the parent comp. Similarly, specific effects applied to the layers could have different results under those conditions. I would bet it is something that trivial. Otherwise your screenshots don't reveal much that would offer clues, unfortunately. Perhaps the flow chart will also reveal some dependency across layers and comps that affects this.
Mylenium